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For example, ''lesion'' to the V5 leads to deficits in perceiving motion and processing of complex stimuli. It contains many neurons selective for the motion of complex visual features (line ends, corners). ''Microstimulation'' of a neuron located in the V5 affects the perception of motion. For example, if one finds a neuron with preference for upward motion in a monkey's V5 and stimulates it with an electrode, then the monkey becomes more likely to report 'upward' motion when presented with stimuli containing 'left' and 'right' as well as 'upward' components.<ref name="BrittenVanWezel">{{cite journal | title=Electrical microstimulation of cortical area MST biases heading perception in monkeys |vauthors=Britten KH, van Wezel RJ | journal=Nature Neuroscience | year=1998 | volume=1 | issue=1 | pages=59–63 | pmid=10195110 | doi=10.1038/259|s2cid=52820462 }}</ref>
There is still much controversy over the exact form of the computations carried out in area MT<ref name="Wilson">{{cite journal | last1 = Wilson | first1 = H.R. | last2 = Ferrera | first2 = V.P. | last3 = Yo | first3 = C. | year = 1992 | title = A psychophysically motivated model for two-dimensional motion perception | journal = Visual Neuroscience | volume = 9 | issue = 1| pages = 79–97 | doi=10.1017/s0952523800006386| pmid = 1633129 | s2cid = 45196189 }}</ref> and some research suggests that feature motion is in fact already available at lower levels of the visual system such as V1.<ref name="Tinsley">{{cite journal | title=The nature of V1 neural responses to 2D moving patterns depends on receptive-field structure in the marmoset monkey | vauthors=Tinsley CJ, Webb BS, Barraclough NE, Vincent CJ, Parker A, Derrington AM | journal=Journal of Neurophysiology | year=2003 | volume=90 | issue=2 | pages=930–7 | pmid=12711710 | doi=10.1152/jn.00708.2002| s2cid=540146}}</ref><ref name="PackBorn">{{cite journal | title=Two-dimensional substructure of stereo and motion interactions in macaque visual cortex |vauthors=Pack CC, Born RT, Livingstone MS | journal=Neuron | year=2003 | volume=37 | issue=3 | pages=525–35 | pmid=12575958 | doi=10.1016/s0896-6273(02)01187-x| doi-access=free }}</ref>
=== Functional organization ===
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